The 76.6 million commuter trips surpassed the old record of 74.9 million recorded in 2008.
The Port Authority says the record ridership follows an investment of more than $1 billion over four years for new cars, station upgrades and the installation of a new signal system to the 104-year-old rail system.
The new cars allow for faster exit and entry.
Harrison and Hoboken stations are undergoing renovations and platforms along the World Trade Center-Newark line will also be extended to accommodate 10-car trains.
Officials say computerized signals along the 43 miles of PATH track will result in fewer delays and better safety.
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